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Dit blog is een onderdeel van EVODISKU
WAT IS DE BEDOELING EN WAAR STAAT DIT BLOG VOOR ****Wie meent dat alles inmiddels wel over de evolutietheorie gezegd is en dat de discussie gesloten kan worden, ziet over het hoofd dat de wetenschap niet stil staat. ***Wie meent dat inmiddels het creationisme definitief het pleit heeft verloren en dat de discussie gesloten kan worden , ziet over het hoofd dat het "creationisme" is geevolueerd ( en zal evolueren ) in nieuwere mimicrytische vormen( meme-complexen ) zoals bijvoorbeeld het ID(C) ***Dit blog is speciaal opgezet om de aktualiteit binnen het evolutie-creationisme debat te volgen en van kommentaren te voorzien ... waartoe de lezers zijn uitgenodigd bij te dragen ... Let echter wél op het volgende : "Je bent een rund als je hier met religie stunt " ....
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27-11-2014 om 08:56 geschreven door doudoune moncler collection hiver 2012 gratuit


29-05-2008
Vogelevolutie
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Terug naar Fossielen Bestand Menu

Vogelevolutie
Over de oorsprong van vogels uit reptielachtige voorouders.

Een artikel van Tim Spaan en J.L.G Arts.


Een tijdlang zijn de meningen verdeelt geweest over welke groep reptielen gestalte heeft gegeven aan de klasse der vogels, een minderheid heeft gedacht dat vogels uit de groep reptielen zijn ontstaan die naar de krokodillen leidt (crocodylomorphen). Veruit de meerderheid heeft echter een geheel andere groep op het oog, namelijk de dinosaurirs. Eerst iets over de andere hypothesen:

De Thecodont hypothese
In 1926 werd Gerhard Heilmanns boek The Origin of Birds gepubliceerd. Dit boek was het resultaat van een zeer uitgebreid onderzoek van de vogelbiologie, inclusief anatomie, embryologie, spierkunde, vleugelonderzoek, paleontologie en vele andere onderwerpen. Heilmann kwam tot de conclusie dat theropode dinosaurirs de meeste overeenkomsten bezaten van alle fossiele groepen tot Archaeopteryx en andere vogels, maar omdat theropode dinosaurussen geen sleutelbeen zouden hebben verwierp hij deze groep als directe voorouders. Hij concludeerde dat de afkomst van vogels moet hebben gelegen bij de meer basale Archosaurirs misschien bij vormen die verbonden zijn aan Ornithosuchus of Euparkeria, welke wel sleutelbeenderen bezaten.
Sinds die tijd zijn er echter wl dinosaurirs ontdekt die sleutelbeenderen hebben, onder andere in de familie der Ceratosaurirs bij Coelophysis en Segisaurus, en een volledig gevormd vorkbeen is recentelijk ontdekt in de theropode Allosaurus, en van tyrannosauriden tot Velociraptor, Oviraptor en Ingenia.
Sinds Heilmann hebben andere paleontologen geprobeerd de thecodont-hypothese te bewijzen, maar zonder veel succes. Geen enkele specifieke basale-archosaurir is kandidaat voor een direct voorouderschap of het dichtstbijzijnde bekende dier tot vogels. De thecodonte oorsprong van vogels staat zwaar onder druk omdat er geen verdediger van de thecodonte oorsprong er in is geslaagd een cladogram te maken die hun hypothese geheel of gedeeltelijk ondersteund.

De Krokodilomorphen hypothese
De aanhangers van deze hypothese denken dat de voorouders van vogels zijn ontstaan in de familie van de krokodillen, maar voordat ware krokodillen zich ontwikkelden. Inderdaad vinden we binnen de groep van sphenosuchische krokodylomorphen, dus buiten de Crocodylia, de meest ontwikkelde vogelkenmerken. A.D. Walker was de voornaamste aanhanger van deze hypothese, hij baseerde zijn bevindingen op zijn gedetailleerde studies van de hersenpan, schedelbeen, oorregio en andere kenmerken in Sphenosuchus, een krokodylomorph uit de Late Jura van Zuid Afrika. Zijn mening werd grotendeels ondersteund door L.D. Martin en zijn studenten, hoewel zij probeerden te bewijzen dat vogels afstammelingen waren van zowel echte krokodillen als krokodylomorphen. Veel van de gevonden overeenkomsten zijn juist geanalyseerd maar bleken veel voorkomend te zijn onder Archosaurirs of gelijktijdig te zijn ontwikkeld in bepaalde krokodillen en vroege vogels (maar niet present in de gehypnotiseerde gezamenlijke voorouder van beide groepen). Wederom kan tot op de dag van vandaag met al het geleverde bewijs geen cladistisch bewijs worden overlegd dat een krokodylomorphische afkomst van vogels ondersteund.

Dinosaurirs
Nu zowel thecodonten als krokodilomorphen als voorouders van vogels niet erg overtuigend meer zijn lijkt de oorspronkelijke conclusie van Gerhard Heilmann, volgens welke de therapode dinosaurirs de meeste overeenkomst vertonen met vogels, de juiste. Met nieuwe ontdekkingen die de laatste tien jaar zijn gedaan is gebleken dat verschillende leden binnen de familie der maniraptora, die wordt onderverdeeld in de families der Dromaeosauridae, Oviraptora, Avialae en Therizinosauridae, grote overeenkomsten vertonen met vogels.
Allereerst zijn er tussen de maniraptora al een groot aantal overeenkomsten in de structuur van het skelet, waarvan dit de meest in het oog springende zijn:

de pubis is van een naar voren gerichte positie veranderd in een naar achteren gerichte positie verlengde armen, handen en tenen grote openingen voor de ogen een speciaal polsgewricht holle beenderen (misschien in verbinding met de longen) 3 vingerige hand, voet met vier tenen waarvan 3 tenen gebruikt worden om te lopen kleinere, achter door pezen verstijfde staart verlengde middenvoetsbeentjes S-vormige lange nek teenganger samengegroeid borstbeen en sleutelbeen enkel, die alleen naar voren en naar achter beweegbaar is schouder die ook zijwaartse en op en neergaande (vleugel) bewegingen toelaatVerder zijn de maniraptora behalve vogels de enige groep dieren die over veren beschikken (Uitgezonderd Sinosauropteryx en Dilong paradoxus) wel dinosaurirs, maar geen maniraptors.), een tiental soorten uit elk van de onderfamilies zijn aangetroffen met fossiele veren, waaronder Microraptor, Beipiaosaurus, Epidendrosaurus en meer. Tevens zijn er maniraptors ontdekt die in een zelfde positie sliepen als moderne vogels, natuurlijk de recentelijk ontdekte Mei, maar ook Sinornithoides is in een dergelijke positie aangetroffen. Tenslotte is ook het feit dat een skelet van de Oviraptor Citipati in een broedhouding over haar nest is gevonden een aanwijzing dat maniraptors niet alleen in skeletstructuur maar ook in gedrag overeen komen met vogels.De conclusie mag duidelijk wezen, dinosaurirs uit de familie maniraptora vertonen veruit de meeste overeenkomst met vogels. In het volgende gedeelte zullen we de bewijzen voor het daadwerkelijke ontstaan van vogels uit maniraptors evalueren. De evolutie van vogels. Het op fylogenetische wijze rangschikken van de voeten van dinosaurirs levert het volgende cladogram op:De rode stippen geven het tijdstip weer waarop de gemeenschappelijke voorouder van vogels en de verschillende families dinosaurirs leefde. Er is een zekere evolutieve ontwikkeling merkbaar, het verbazende is dat exact dezelfde ontwikkeling plaatsvind in de embryo's van moderne vogels.Een andere aanwijzing is de staart. Je ziet dat hoe dichter de dinosaurussen bij vogels staan, hoe minder staartwervels er zijn. Om tot Aves gerekend te kunnen worden mag de staart niet langer zijn dan 25-26 wervels. Bij primitieve theropoden (ceratosauria) zien we zo'n 50 staartwervels en bij coelurosauria niet meer dan 40. Als we verder bij de vogels komen zien we dat de staart minder beweeglijk wordt. Het zogenaamde transition point komt steeds dichter bij het begin van de staart te liggen. Dit punt geeft de overgang aan van de beweeglijke staartwervels naar de onbeweeglijke staartwervels. Dit komt overeen met het eindpunt van de chevronbeentjes, dit zijn de doornachtige uitsteeksels onder de staartwervels. Bij de wat modernere vogels vergroeien de staarwervels geheel en vormen de voor vogels zo karakteristieke Pygostyle. Bij embryos bestaat de staart trouwens nog uit losse wervels die pas later vergroeien.Tenslotte de heupen, Dinosaurirs worden op basis van hun heupen in 2 groepen verdeeld, de Saurischia (Hagedis-heupen) en Ornithischia (Vogel-heupen). Nu behoren de voorouders van de vogels niet tot de Ornithischia maar tot de Saurischia. Echter zoals je kunt zien verplaatst de (met groen aangegeven) pubis (schaambeen) zich telkens wat naar achteren om uiteindelijk een vogelachtige heup te vormen.Dit zijn dus de voornaamste evolutieve ontwikkelingen die zij

29-05-2008 om 12:47 geschreven door Tsjok45


18-09-2007
Gevederde dino's
Coelurosauria
  Tugulusaurus (may or may not have been feathered)
 
 
Tyrannosauroidea (primitive feathers)
        Dilong paradoxus (feather impressions)
        Tyrannosaurus (scale impressions, may have lost some feathers)
   
   
Compsognathidae (primitive, two-branched feathers)
                   Sinosauropteryx (feather impressions)
                   Juravenator (scale impressions, may have lost some feathers)
                   Compsognathus (feathers possible)
   
   
Maniraptoriformes
         Ornithomimosauria (feathers likely)
        
Alvarezsauridae (feathers with a central vane)
                    Shuvuuia (feathers preserved in three dimensions)
                    Mononykus (feathers extremely likely)
        
        
Maniraptora (flight feathers and down-like contour feathers)
              Yixianosaurus (feather impressions)
             
              Oviraptoriformes (vaned, plumaceous feathers)
                  
                  
Oviraptorosauria
                        Protarchaeopteryx (feather impressions)
                        Caudipteryx (feather impressions)
                        Avimimus (feathers extremely likely)
                       
Oviraptoridae (feathers extremely likely, skeletal feather adaptations, behavioral clues in nesting Citipati)
                  
                  
Therizinosauroidea
                          Beipiaosaurus (feather impressions)
             
              Paraves (flight feathers and down-like contour feathers)
                    Pedopenna (feather impressions)
                   
                   
Deinonychosauria (asymmetrical feathers)
                            
Troodontidae
                                   Jinfengopteryx (feather impressions)
                                   Troodon (feathers likely)
                            
                            
Dromaeosauridae
                                    Rahonavis (skeletal feather/flight adaptations)
                                    Cryptovolans (feather impressions)
                                    Microraptor (feather impressions)
                                    Sinornithosaurus (feather impressions)
                                    Velociraptor (feathers extremely likely)
                   
                    Epidendrosaurus (feather impressions)
                   
                    Aves
(asymmetrical flight feathers)



Sinosauropteryx (1996)    Sinosauropteryx<?xml:namespace prefix = o ns = "urn:schemas-microsoft-com:office:office" />


    
18-09-2007 om 21:10 geschreven door Tsjok45


13-09-2007
A Basal Dromaeosaurid and Size Evolution Preceding Avian Flight

Science 7 September 2007:
Vol. 317. no. 5843, pp. 1378 - 1381
DOI: 10.1126/science.1144066

 

Reports

A Basal Dromaeosaurid and Size Evolution Preceding Avian Flight

Alan H. Turner,1*Diego Pol,2Julia A. Clarke,3,4,1Gregory M. Erickson,5Mark A. Norell1

Fossil evidence for changes in dinosaurs near the lineage leadingto birds and the origin of flight has been sparse. A dinosaurfrom Mongolia represents the basal divergence within Dromaeosauridae.The taxon's small body size and phylogenetic position implythat extreme miniaturization was ancestral for Paraves (theclade including Avialae, Troodontidae, and Dromaeosauridae),phylogenetically earlier than where flight evolution is stronglyinferred. In contrast to the sustained small body sizes amongavialans throughout the Cretaceous Period, the two dinosaurianlineages most closely related to birds, dromaeosaurids and troodontids,underwent four independent events of gigantism, and in somelineages size increased by nearly three orders of magnitude.Thus, change in theropod body size leading to flight's originwas not unidirectional.

1 Division of Paleontology, American Museum of Natural History, Central Park West at 79th Street, New York, NY 100245192, USA.
2 CONICET, Museo Paleontológico Egidio Feruglio, Avenida Fontana 140, (9100) Trelew, Argentina.
3 Department of Marine, Earth and Atmospheric Sciences, North Carolina State University, Campus Box 8208, Raleigh, NC 276958298, USA.
4 Division of Paleontology, North Carolina Museum of Natural Sciences, 11 West Jones Street, Raleigh, NC 276011029, USA.
5 Department of Biological Sciences, Florida State University, Dewey Street and Palmetto Drive, Tallahassee, FL 323061100, USA.

* To whom correspondence should be addressed. E-mail: turner@amnh.org

Which nonflying maniraptoran dinosaurs are the closest relativesto birds (Avialae) has been debated (15). Dromaeosauridsand troodontids are the two clades consistently found to bemost closely related to avialans (18). Discoveries ofthese dinosaurs, which illuminate the features ancestrally presentin the first flighted theropods, have remained rare. Here wereport a basal dromaeosaurid theropod: Theropoda Marsh, 1884;Maniraptora Gauthier, 1986; Paraves Sereno, 1997; DromaeosauridaeMatthew and Brown, 1922; Mahakala omnogovae, new taxon. Thenew taxon is small (~70 cm long) and possesses features absentin other dromaeosaurids but shared with early troodontids andavialans.

Holotype. Specimen number IGM (Mongolian Institute of Geology,Ulanbaatar) 100/1033, a partial skull and postcranial skeleton(Figs. 1 and 2).


Figure 1 Fig. 1. Mahakala omnogovae IGM 100/1033, holotype. (A) Skull in occipital view. (B) Braincase in left lateral view. (C) Sacrum and partial right leg in ventral view. (D) Frontal in dorsal (left) and ventral (right) views. (E) Axisvertebrain left lateral view. Scale bars, 5 mm in (A), (B), (D), and E) and 1 cm in (C). Abbreviations are as follows: cav, caudal vertebra; ctr, caudal tympanic recess; dtr, dorsal tympanic recess; ep, epipophysis; f.l, lacrimal facet; f.po, postorbital facet; fm, foramen magnum; mt, metatarsus; oc, occipitalcondyle; od, odontoid; pap, paroccipital process; pf, pneumatic foramen; ph, phalanx; prz, prezygapophysis; q.pr, contactsurface on prootic for quadrate; q, quadrate; ti, tibia; tl, tectal lobe; sac, sacrum; v.o, occipital vein track. [View Larger Version of this Image (40K GIF file)]
 

Figure 2 Fig. 2. Mahakala omnogovae IGM 100/1033, holotype. (A) Rightulna in lateral (right) and medial (left) views. (B) Ilium in medial (top) and lateral (bottom) views. (C) Femurinposterior (left) and lateral (right) views. (D) Tibia in anterior view. (E) Left metatarsus in anterior view. (F) Right raptorial claw. (G) Midcaudal vertebrae. Scale bars, 1 cm in (B) to (E) and 5 mm in (A), (F), and (G). Abbreviations are as follows: aa, ascending process of astragalus; as, astragalus; bf, brevis fossa; brs, brevisshelf; bs, biccipital scar; ca, calcaneum; cc, cnemial crest; ch, chevron; fc, fibular crest; fi, fibula; gtr, greater trochanter; lr, lateralridge; lc, lateral crest; mco, medial condyle; mt, metatarsal; obr, oblique ridge; pat, posterior antitrochanter; prz, prezygapophysis; pt, posterior trochanter; ts, trochanteric shelf. [View Larger Version of this Image (45K GIF file)]
 

Etymology. "Mahakala," Sanskrit for one of the eight protectordeities (dharmapalas) in Tibetan Buddhism. The specific epithetrefers to the southern Gobi provenance of this taxon.

Locality and horizon. The Tugrugyin Member of the DjadokhtaFormation (Campanian) (9, 10), Tugrugyin Shireh, Ömnögov,Mongolia (10, 11).

Diagnosis. A small paravian diagnosed by the following combinationof characters (autapomorphies are noted by *): a strongly compressedand anteroposteriorly broad ulna tapering posteriorly to a narrowedge*; an elongate lateral crest on the posterodistal part ofthe femur*; anterior caudal vertebrae with subhorizontal, laterallydirected prezygapophyses*; a prominent supratrochanteric process;and the absence of a cuppedicus fossa.

Estimated at 70 cm long, Mahakala is similar in size to thebasal avialan Archaeopteryx and basal members of other maniraptoranclades such as the oviraptorosaur Caudipteryx and the troodontidMei long. The specimen is a young adult or near adult, basedon the degree of neurocentral and astragalocalcaneal fusion,braincase coossification, and histological analysis (fig. S4).Thus, it can be distinguished from the contemporaneous Archaeornithoides,which is of similar size but is a juvenile (12).

The braincase, quadrate, and frontals are well preserved. Unlikedromaeosaurids but similar to troodontids such as Sinovenator(7) and Mei (8), the frontals are dorsoventrally vaulted andthe interorbital region is narrow, indicating proportionallylarge orbits. The anterolateral corner of the frontal lacksthe articulation notch present in other dromaeosaurids. Thefrontals transition smoothly from the orbital margin to thepostorbital processes as in troodontids (13), but unlike theabrupt transition and sharply demarcated postorbital processesof dromaeosaurids. The supratemporal fossa margin is weaklycurved, not sinuous as in all other dromaeosaurids except Tsaagan(5) and Dromaeosaurus [AMNH (American Museum of Natural History)FR 5356]. The quadrate is incipiently bistylic, unlike the single-headedball-shaped process in other dromaeosaurids. A depression onthe prootic may correspond to a secondary articulation surfacefor the quadrate. This depression corresponds topographicallyto the braincase articulation facet in birds and alvarezsauridsbut is also present in the troodontid Byronosaurus (14). Thelateral braincase wall lacks any indication of a well-developedotosphenoidal crest like that in troodontids (14). The paroccipitalprocesses are relatively short and distally twisted rostrolaterallyas in other dromaeosaurids.

The axis bears a single pneumatic opening and has small epipophysesthat do not overhang the postzygapophyses. The sacrum comprisessix apneumatic, coossified centra as in Rahonavis and maturespecimens of Velociraptor (IGM 100/986 and IGM 100/985). Thefused neural arches of the posterior sacral vertebrae form abony lamina as in other dromaeosaurids. The tail is long asin basal avialans (such as Archaeopteryx and Jeholornis), basaltroodontids (such as Jinfengopteryx and Mei), and other dromaeosaurids.The transition point occurs between caudals (Cd's) 11 and 12and is more posteriorly located than in Rahonavis (proximalto Cd 9) or Velociraptor (proximal to Cd 10). The distal caudalpostzygapophyses are smaller than the prezygapophyses. As into other basal paravians, the postzygapophyses do not exceedthe posterior margin of the vertebral centra. The lateral surfaceof the proximal caudals bears a low ridge similar to that inBuitreraptor (4) and Rahonavis [UA (University of Antananarivo)8656]. The chevrons in Mahakala are platelike as in many derivedcoelurosaurs.

The scapula is narrow and straplike and has a strongly compressedovoid cross section. The preserved portion of the incompletehumerus suggests that the entire humerus was reduced in contrastto the condition of most coelurosaurs. The ulna is distinctlybowed as in most maniraptorans (1) but is strongly compressedand possesses a small biceps tubercle. The distal region ofthe radius is expanded and flattened as in paravians (11). Asemilunate carpal covers the proximal surfaces of metacarpalsI and II; a plesiomorphic conformation lost in most avialans.

The ilium is dolichoiliac. A prominent supratrochanteric processis present as in Unenlagia, Rahonavis, and many avialans. Thebrevis shelf is triangular and does not extend laterally asin some other basal dromaeosaurids. No antiliac shelf is present,therefore Mahakala lacks a defined cuppedicus fossaanabsence unique within dromaeosaurids but characteristic of avialanssuch as Apsaravis (15) and Yixianornis (16).

The femur is anteriorly bowed. The lesser trochanter is welldeveloped, and its anterior edge is continuous with the greatertrochanter. The fourth trochanter is present as a smooth andweakly developed ridge. Unlike Velociraptor (IGM 100/986), thelateral ridge is poorly developed, and the moundlike trochantericshelf is proximodistally elongated and closely connected tothe posterior trochanter. A prominent crest extends from thedistal third of the shaft to the ectocondylar tubercle. Thetibia is longer than the femur and possesses a single cnemialcrest. The lateral surface of the calcaneum is distinctly concaveand lacks the notch for the articulation with the distal fibulathat is present in dromaeosaurids and other nonavian theropods.This condition is shared with Rahonavis, basal avialans (2),and derived alvarezsaurids. Unlike most troodontids and microraptorines,but similar to Archaeopteryx and derived dromaeosaurids, thefoot of Mahakala exhibits the plesiomorphic unconstricted conditionfor metatarsal III, further indicating that this avian traitmay be the primitive condition for paravians. The distal endof metatarsal II is composed of an asymmetrical ginglymoid articularsurface and phalanx II-2 has a well-developed proximal heeland hypertrophied ginglymoid trochlea. This suite of charactersis present only in dromaeosaurids.

Phylogenetic analysis identifies Mahakala as a basal dromaeosauridand supports paravian monophyly with birds (Avialae) as thesister group to a monophyletic Deinonychosauria (Dromaeosauridae+ Troodontidae) (Fig. 3 and fig. S1). Although discovered inrelatively young Cretaceous deposits, the basal position ofMahakala has several implications regarding our understandingof the early history of deinonychosaurians (17). First, Shanagfrom the Early Cretaceous of Mongolia (18) nests within thepurported Gondwanan lineage of dromaeosaurids, Unenlagiinae.This topology complicates recently proposed vicariance-drivenorigin hypotheses for these groups (4, 19). Second, these dinosaursare united with Jehol microraptorines (Microraptor, Graciliraptor,and Sinornithosaurus) to form the sister group to derived dromaeosauridsfrom Laurasia (velociraptorines and allied forms). Third, thepurported avialan Jinfengopteryx (20) is a troodontid. Jinfengopteryxhas feathers; it thus demonstrates the presence of feathersof modern aspect in a troodontid.


Figure 3 Fig. 3. Phylogeny and body size change within paravian theropods. A temporally calibrated cladogram depicting the phylogenetic position of Mahakala and paravian body size through time and across phylogeny is shown. Characters uniting Mahakala with other dromaeosaurids include the absence of an accessory tympanic recess dorsal to the crista interfenestralis, and elongate paroccipital process with parallel dorsal and ventral edges that twist rostrolaterally distally, and the presence of a distinct ginglymus on the distal end of metatarsal II (17). Silhouettes are to scale, illustrating the relative magnitude of body size differences. Left-facing silhouettes near open circles show reconstructed ancestral body sizes. Ancestral paravian body size is estimated to be 600 to 700 g and 64 to 70 cm long (17). The ancestral deinonychosaur, troodontid, and dromaeosaurid body size is estimated at ~700 g. Large numbers (1, 2, 3, and 4) indicate the four major body increase trends in Deinonychosauria. See the supporting online material for further ancestral body size reconstruction data. Ma, Maastrichtian; Ca, Campanian; Sa, Santonian; Co, Coniacian; Tu, Turonian; Ce, Cenomanian; Ab, Albian; Ap, Aptian; Bar, Barremian; Hau, Hauterivian; Va, Valanginian; Ber, Berriasian; Ti, Tithonian; Ki, Kimmeridgian. Ma, million years ago. [View Larger Version of this Image (35K GIF file)]
 

Decrease in body size is a trend in coelurosaurs (3, 8, 21)and is thought to have played an important role in the originof birds and flight (6, 11, 2224). Dromaeosaurids andother coelurosaurs, however, may have undergone clade-specificincreases in body size (8, 25). Testing these trends requiresempirical size reconstructions for each node of the coelurosaurtree. We estimated ancestral body sizes for each internal node(ancestral node) using body mass estimates from femoral lengthmeasurements. These data were treated as a continuous additivetrait and optimized across the phylogeny (16).

Our analysis (fig. S2) indicates that small body size was nota derived condition at Archaeopteryx or Avialae, where flightevolution in theropods is currently inferred. The ancestraldromaeosaurid, troodontid, and deinonychosaurian are reconstructedas small, each with a body mass around 700 g (Fig. 3). The basalmembers of these lineages are the same size as the early avialanJeholornis. Additionally, our results indicate that deinonychosaursunderwent four parallel trends of body size increase. Threeof these events occurred within Dromaeosauridae: Deinonychusincreased in size by more than two orders of magnitude, as didUnenlagia, and the Achillobator + Utahraptor clade increasedby three orders of magnitude. A single trend of body size increasewas observed in troodontid body size. These events were contemporaneouswith a decrease in avialan body sizes. Our analysis impliesthat the ancestral paravian had a body size of 600 to 700 gand was ~65 cm long, roughly the size of the largest specimensof Archaeopteryx or Sapeornis and entailing the size range reconstructedfor basal deinonychosaurs. Thus, miniaturization preceded theavialan node and the origin of flight, and as a result, hypothesesrelating ontogenetic or metabolic controls on miniaturizationto flight origin in theropods must be equally capable of explainingthe size reduction within ancestral paravians and the iterativetrends of size increase in deinonychosaurs.


References and Notes

1. J. Gauthier, Mem. Calif. Acad. Sci. 8, 1 (1986). 2. C. A. Forster, S. D. Sampson, L. M. Chiappe, D. W. Krause, Science 279, 1915 (1998). [CrossRef] [ISI] [Medline] 3. P. C. Sereno, Science 284, 2137 (1999). [CrossRef] [ISI] [Medline] 4. P. J. Makovicky, S. Apesteguía, F. L. Agnolin, Nature 437, 1007 (2005). [CrossRef] [Medline] 5. M. A. Norell et al., Am. Mus. Novit. 3545, 1 (2006). [CrossRef] 6. X. Xu, Z. Zhou, X.-l. Wang, Nature 408, 705 (2000). [CrossRef] 7. X. Xu, M. A. Norell, X.-l. Wang, P. J. Makovicky, X.-C. Wu, Nature 415, 780 (2002). 8. M. A. Norell, X. Xu, Nature 431, 838 (2004). [CrossRef] 9. Z. Kielan-Jaworowska, R. Barsbold, Palaeontol. Pol. 27, 5 (1972). 10. D. Dashzeveg et al., Am. Mus. Novit. 3498, 1 (2005). [CrossRef] 11. M. A. Norell, P. J. Makovicky, Am. Mus. Novit. 3282, 1 (1999). 12. A. Elzanowski, P. Wellnhofer, Am. J. Sci. 293, 235 (1993).[Abstract/Free Full Text] 13. P. J. Makovicky, M. A. Norell, in The Dinosauria, D. B. Weishampel, P. Dodson, H. Osmólska, Eds. (Univ. of California Press, Berkeley, CA, ed. 2, 2004), pp. 184195. 14. P. J. Makovicky, M. A. Norell, J. M. Clark, T. Rowe, Am. Mus. Novit. 3402, 1 (2003). [CrossRef] 15. M. A. Norell, J. A. Clarke, Nature 409, 181 (2001). [CrossRef] 16. J. A. Clarke, Z. Zhou, F. Zhang, J. Anat. 208, 287 (2006). [CrossRef] [ISI] [Medline] 17. Materials and methods are available as supporting material on Science Online. 18. A. H. Turner, S. H. Hwang, M. A. Norell, Am. Mus. Novit. 3557, 1 (2007). [CrossRef] 19. F. E. Novas, D. Pol, Nature 433, 858 (2005). [CrossRef] [Medline] 20. Q. Ji et al., Geol. Bull. China 24, 197 (2005). 21. M. T. Carrano, in Amniote Paleobiology: Perspectives on the Evolution of Mammals, Birds, and Reptiles, M. T. Carrano, T. J. Gaudin, R. W. Blob, J. R. Wible, Eds. (Univ. of Chicago Press, Chicago, 2006), pp. 225268. 22. Z. Zhou, Naturwissenschaften 91, 455 (2004). [CrossRef] [ISI] [Medline] 23. E. Buffetaut et al., Naturwissenschaften 92, 477 (2005). [CrossRef] [ISI] [Medline] 24. K. Padian, A. J. de Ricqlés, J. R. Horner, Nature 412, 405 (2001). [CrossRef] 25. X. Xu, Q. Tan, X. Zhao, L. Tan, Nature 447, 844 (2007). [Medline] 26. We thank the field crew of the 1993 field season for their work; X. Xu, Z. Zhou, C. Forster, and D. Krause for specimen access; P. Makovicky, N. Smith, J. Conrad, A. Balanoff, G. Bever, R. Irmis, and S. Nesbitt for discussions; M. Ellison for photographs; and B. Amaral, A. Davidson, and A. Balcarcel for preparation. Support was provided by NSF through a Doctoral Dissertation Improvement Grant (DEB 0608003, presented to A.H.T. and M.A.N.); grant ATOL 0228693 (presented to M.A.N.); the Program in Geoscience, Division of Earth Sciences (grant EAR 0207744, presented to G.M.E. and M.A.N.); and the Division of Biological Infrastructure, Program in Biological Databases and Information (grant DBI 0446224, presented to G.M.E.). Additional support was provided to A.H.T. by the American Museum of Natural History and Columbia University.
Supporting Online Material

www.sciencemag.org/cgi/content/full/317/5843/1378/DC1

SOM Text

Figs. S1 to S5

References


Received for publication 20 April 2007. Accepted for publication 30 July 2007.

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    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen. PLACENTALE ZOOGDIEREN STAMBOOM CONTROVERSE
     
  • Een uitgekomen voorspelling
  • Wie oren heeft , die hore
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.Eritherium azzouzorum
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.LJOEBA
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.Darwinius masillae
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.VLIEGENDE KATTEN   ?
    Vleermuizen
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.ALWEER EEN BELANGRIJK STUK UIT DE LEGPUZZEL
    Zee-zoogdieren ;
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.Van de wal in de visgronden  Indohyus / walvisevolutie
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.Zeehonden-evolutie

    amfibieen 


    Vissen
    Overgang  naar tetrapoda

     
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.Opnieuw Tiktaalik
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.Ventastega

    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.KIEKEBOE BOE BOE (oude tetrapoden )
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.De niet-zo-missing links gallerij (1) 

    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.KWASTEN
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.KWASTEN (2)

    Longvissen
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen. HELP TE WEINIG ZUURSTOF


    Arthropoda
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.trilobieten

    Weekdieren
    OCTOPUSSY


    d) Geologie
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.ALH 84001
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.CHICXULUB
    TIPS

    *Blauwe
    en groene ( =nederlandse)teksten zijn meestal  aanklikbare links 
    *Engelse Wikipedia teksten  verwijzen   in de  linkerkolom naar  verschillende  niet-engelse versies van het   wikiartikel
    *Blauwe teksten tusssen ""  ,  zijn voornamelijk  ( gedeeltelijke)citaten afkomstig van  mensen met andere  meningen


      







    Links naar  Gespecialiseerde  Verenigingen en sites


    Fossielnet
    http://www.fossiel.net/

    Arachnologie  
    ARABEL

    http://www.arabel.ugent.be/

    Myrmycologie
    Mierenwerkgroep * Polyergus *
    http://formicidae.be/

    Entomologie
    Jean-Luc Renneson
    http://users.swing.be/entomologie/
    http://users.swing.be/entomologie/Vespidae%20of%20the%20World1.htm


    Three of life  projects 



     "The Tree of Life".
     
     
  • Tree of Life Web Project
  • Green Tree of Life at Berkeley
  • Fungal Tree of Life Project
  • Beetle Tree of Life project
  • Fly Tree of Life project
  • Mammal Tree of Life project
  • Cypriniformes Tree of Life project
  • Liverwort Tree of Life Project
  • Early Bird Tree of Life project
  • Early Bird Tree of Life project
  • Angiosperm Tree of Life project
  • Cnidaria Tree of Life Project
  • Decapoda Tree of Life Project

    LepTree.net,
    the Tree of Life project on Lepidoptera.


    http://www.rebeccashapley.com/cipres/telescoping.htm
    The Tree of Life blog
    Tree of Life at wikipedia.
  • Zoeken met Google


    Zoekmachine ;
     


    doorzoekt 4-sites, inclusief:
    http://tsjok45.multiply.com/photos
    http://anticreato.multiply.com/,
    http://evodisku.multiply.com/,
    http://www.bloggen.be/evodisku/ ,




      


    site search by freefind advanced

    Tomaso schrijft over      

    Evolutie /  Charles Darwin aan de basis:
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.SELECTIE NATUURLIJK
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.DE LEVENSBOOM
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.UITSTERVEN
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.DEEP TIME
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.Biogeografie
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.SEKSUELE SELECTIE
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.Al jaren lang samen CO-EVOLUTIE
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.De economie van de natuur
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.Geleidelijke veranderingen

    Unintelligent design:

    Blog Entry
    Ons gebit , De appendix,  Geboorte, 
    De weg van het zaad , Nervus  Laryngeus Recurrens,
    DNA , 75 mistakes , Het boek 

    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.Unintelligent design (1): Rechtop lopen
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.Tomaso over het oog , oogspieren ,
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.Unintelligent Design (5): ZWEETVOETEN 
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.Unintelligent design (6): Vitamine C 
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.Unintelligent design (8): Kuitspieren
    DARWINJAAR

    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.HET GELIJK VAN DARWIN

    Darwin is GEEN Heilige Sinterklaas
    2009 Année Darwin
    De eerste pasjes ?
    Darwinjaar trekt zich op gang I
    Darwinjaar trekt zich op gang II


    foto





















    Nederlandse  Blog-zwerm  /een selectie
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.me.myself.and.I.

    De Stoet

    Tsjok45    zie ook  MULTIPLY evodisku
    http://evodisku.multiply.com/tag/darwinjaar
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.PETER LOUTER
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.STRIPMAN
    Theodorus

    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.RAMIREZI
    http://ramireziblog.blogspot.com/

    Cees Chamuleau
    Zoe van Zaal
    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.MIEKE ROTH

    100 Woorden

    Klik hier om een link te hebben waarmee u dit artikel later terug kunt lezen.Jeroen de Baaij
    Ragfijn
    Annelies
    Satuka

    Qabouter 1 Qabouter 2


    Pierra
    Johan H.v.D.
    Theo E. Korthals Altes

    Meneer Opinie 1 Meneer Opinie 2 Meneer Opinie 3

    Tomaso Tomaso 2

    Laila (??)
    Linda Morgan
    Wilma
    Kees Smit
    Henk van Leuken
    Iris
    Vroems
    Erik de Groot
    Ely
    Mephisto

    Thera

    Bart
    Rein John
    Martijn van Calmthout Martijn van Calmthout 2
    Draakhond
    Jeg Synes
    Jim Hasenaar
    Kokopelli
    Jos Goedmakers
    Fleur Frenkel Frank
    London Calling
    Maria Trepp
    Johan H. van D. 2
    Aad Verbaast
    Landheha
    Ina Dijstelberge


    sterk aanbevolen 
     
    Terrence :
    http://boomdeslevens.blogspot.com/    
    een voortzetting van het
    vroegere -->
    http://www.vkblog.nl/blog/125817/Neomuran_hypothesis

      
    De sublieme   
    PIERRA
    (uiteraard ook sterk aanbevolen )


    http://ascendenza.wordpress.com/2011/01/08/welkom-op-mijn-nieuwe-blog/
      http://www.vkblog.nl/blog/95678/Op_zoek_naar_de_klepel
     



    Tsjok45 photo site  MULTIPLY



    MULTIPLY evodisku 

    Blog Entry





     


     


    ANTI-CREATO   MULTIPLY



    Blog Entry INHOUD ANTI CREATO


    LINKS  NAAR VERKLARENDE  NOTEN
    (In verschillende talen )

  • INDEX   NOTES  : 
    http://evodisku.multiply.com/tag/inhoud%20%20notes

  • T -->

    Blog tegen de wet? Klik hier.
    Gratis blog op https://www.bloggen.be - Bloggen.be, eenvoudig, gratis en snel jouw eigen blog!